Dorcatherium from the dhok bun ameer khatoon, chinji formation, lower siwalik hills of Pakistan
Paper Details
Dorcatherium from the dhok bun ameer khatoon, chinji formation, lower siwalik hills of Pakistan
Abstract
Several new fossil specimens of Dorcatherium have been found and described from the Chinji Formation near Dhok Bun Ameer Khatoon (DBAK). The material comprises upper and lower dentitions and presents the data which is new, also it provide us further information about the distribution of two species Dorcatherium majus and Dorcatherium minus from the Lower Siwalik locality. The fossil site has been dated to approximately 14.2 Ma – 9.5 Ma and coincides with the divergence of different mammalian genera. It has well-exposed Chinji and Nagri formations. The genera recovered from this site are mostly the same as in the overlying younger Dhok Pathan Formation of the Siwaliks. Size differences between these and previously described specimens are potentially important for our understanding of intraspecific variation in mammalian lineages. This paper therefore describes the newly recovered specimens, assigns them to species level, and then goes on to examine size differences between these and the type material.
Agnarsson I, Maycollado LJ. 2008. The phylogeny of Cetartiodactyla: The importance of dense taxon sampling, missing data and the remarkable promise of cytochrome b to provide reliable species level phylogenies. Molecular Phylogenetics and Evolution 48, 964–985. http://dx.doi.org/10.1016/j.ympev.2008.05.046
Barry J, Morgan M, Flynn L, Pilbeam D, Behrensmeyer AK, Raza SM, Khan I, Badgely C, Hicks J, Kelley J. 2002. Faunal and Environmental change in the Late Miocene Siwaliks of northern Pakistan. Palaeobiology 28,1–72.
Barry JC, Jhonson MN, Raza SM, Jacobs LL. 1985. Neogene mammalian faunal change in Southern Asia: Correlations with climatic, tectonic, and eustatic events. Geology 13, 637–640.
Barry JC, Morgan ME, Weinkler AJ, Flynn LJ, Lindsay EH, Jacobs LL, Pilbeam D. 1991. Faunal interchange and Miocene terrestrial vertebrates of southern Asia. Paleobiology 17, 231–245.
Behrensmeyer A, Barry J. 2005. Biostratigraphic Surveys in the Siwaliks of Pakistan. A Method for Standardized Surface Sampling of the Vertebrate Fossil Record. Palaeontology Electronica 8(1), 1–24.
Behrensmeyer AK, Deino AL, Hill A, Kingston JD, Saunders JJ. 2002. Geology and geochronology of the middle Miocene Kipsaramon site complex, Muruyur beds, Tugen Hills, Kenya. Journal of Human Evolution 42, 1–38.
Bhatti ZH. 2005. Taxonomy, Evolutionary History & Biogeography of the Siwalik Giraffids. Ph. D dissertation, University of the Punjab, Pakistan.
Cheema IU. 2003. Phylogney and evolution of Neogene murine Rodents from the Potwar Platean of Pakistan and Azad Kashmir with special emphasis on zoogeographic diversification and straigraphic. Ph.D. Thesis, University of the Punjab Lahore.
Colbert EH. 1935. Siwalik mammals in the American Museum of Natural History. Transition American Philosophy Society 26, 1–401.
Farooq U, Khan MA, Akhtar M, Khan AM, Ali Z. 2007a. Dorcatherium minus from the Siwaliks, Pakistan Journal of Animal and Plant Science 17, 3–4.
Farooq U, Khan MA, Akhtar M, Khan AM. 2007c. Dorcabune anthracotherioides (Artiodactyla, Ruminantia, Tragulidae) from Hasnot, the Middle Siwaliks, Pakistan. Pakistan Journal of Zoology 39(6), 353–360.
Farooq U, Khan MA, Akhtar M, Khan AM. 2008. Lower dentition of Dorcatherium majus (Tragulidae, Mammalia) in the Lower and Middle Siwaliks (Miocene) of Pakistan. Turkish Journal Zoology 32, 91–98.
Farooq U, Khan MA, Akhtar M. 2007. Dorcatherium majus, a study of upper dentition from the Lower and Middle Siwaliks of Pakistan. Journal of Applied Science 7(9), 1299–1303.
Farooq U, Khan MA, Akhtar M. 2007b. Dorcatherium majus, a study of upper dentition from the Lower and Middle Siwaliks of Pakistan. Journal of Applied Sciences 7(9), 1299–1303.
Farooq U, Khan MA, Akhtar M. 2007d. Dorcabune nagrii (Ruminantia, Tragulidae) from the Upper Part of the Middle Siwaliks. Journal of Applied Sciences 7(10), 1428–1431.
Farooq U. 2006. Studies of evolutionary trends in dentition of the Siwalik Tragulids. Ph.D. Dissertation University of the Punjab, Pakistan.
Gaur R. 1992. On Dorcatherium nagrii (Tragulidae, Mammalia) with a review of Siwalik tragulids. Riv. Ital. Paleontol. Stratigr 98, 353–370.
Gentry A, Rossner G, Heizmann. 1999. Suborder ruminantia in Rossner, G, E., Hessing, K, (Eds). The Miocene land Mammals of Europe. Munchen, Verlag dr. Fpfeil 225–258.
Gentry AW, Hooker JJ. 1988. The phylogeny of Artiodactyla; pp: 235-272. In Benton, M.J. (ed.) The Phylogeny of the Tetrapods. Vol. 2. Mammals. Clarendon Press, Oxford. Guan J. 1988. The Miocene strata and mammals from Tongxin, Ningxia and Guanghe, Gansu. Memoirs of Beijing Natural History Museum 42, 1–21.
Gentry AW. 1978. Tragulidae and Camelidae. In: Evolution of African Mammals (V.S. Maglio and H.B.S. Cooke, eds.) pp. 536–539. Harvard University Press Cambridge Mass.
Gentry AW. 1999. Fossil Pecorans from the Baynunah Formation, Emirate of Abu Dhabi, United Arab Emirates. In: Whybrow P.J. Hill A.,eds. Fossil Vertebrates of Arabia. Yal University Press, New Haven 290–316.
Gentry AW. 2000. The Ruminant Radiation. (eds. E.S. Vrba and G.B. Schaller), Antelopes, deer and relatives: fossil record, behavioural ecology, systematics and conservation. Yale University Press, New Haven 11-25.
Geraads D, Werdelin L, Sanders WJ. 2010. Cenozoic Mammals of Africa University of California Press.
Ghaffar A. 2010. GIS and paleontology of Dhok Bun Ameer Khatoon fossil site, Pakistan. Pakistan Journal of Science 62.
Hill A, Leakey MG, Kingston JD, Ward S. 2002. New cercopithecoids and a hominoid from 12.5 Ma in Tugen Hills succession, Kenya. Journal of Human Evolution 42, 75–93.
Janis CM. 1984. Tragulids as Living Fossils. In: Eldredge, N. and S.M. Stanley (Eds) Living Fossils. New York, Berlin, Heidelberg, Tokyo: Casebooks in Earth Sciences, pp: 87–94.
Johnson GD, Zeitler P, Naeser CW, Johnson NM, Summers DM, Frost CD, Opdyke ND, Tahirkheli RAK. 1982. The occurrence and fission-track ages of Late Neogene and Quaternary volcanic sediments, Siwalik group, northern Pakistan. Paleogeography Paleoclimatology Paleoecology 37, 63–93.
Kaup JJ. 1833. Mitteilungen an Professor Bronn. Neues Jahrbuch fürMineralogie, Geognosie, Geologie und Petrefaktenkunde Jahrgang, Pp 419–420.
Kay RNB. 1987. The comparative anatomy and physiology of digestion in tragulids and cervids, and its relation to food intake. In: C. M. Wemmer (ed.), Biology and Management of the Cervidae, Research Symposia of the National Zoological Park 214–222.
Khan MA, Akhtar M, Ghaffar A, Iqbal M, Khan AM, Farooq U. 2008. Early ruminants from Dhok Bin Mir Khatoon (Chakwal, Punjab, Pakistan): Systematics, Biosratigraphy and Paleoecology. Pakistan journal of Zoology 40(6), 457–463.
Khan MA, Akhtar M, Rohi G, Iqbal M, Samiullah K. 2011. Sivaceros gradiens Pilgrim 1937 (Mammalia, Bovidae, Boselaphini) from the Lower Siwaliks of Dhok Bun Amir Khatoon, Chakwal, Pakistan: Systematics and Biostratigraphy. Turkish Journal of Zoology 35(2), 281–286. http://dx.doi.org/10.3906/zoo-0812-18
Khan MA, Akhtar M. 2011. Dorcatherium cf. nagrii from the Chinji Type Locality (Chakwal, northern Pakistan) of the Chinji Formation, Lower Siwaliks, Pakistan. Pakistan Journal of Zoology 43(6), 1101–1109.
Khan MA, Akhter M. 2013. Tragulidae (Artiodactyla, Ruminantia) from the Middle Miocene Chinji Formation of Pakistan. Turkish Journal of Earth Science 22, 339–353. http://dx.doi.org/10.3906/yer-1106-6
Khan MA, Butt SS, Khan AM, Akhtar M. 2010. A new collection of Giraffokeryx punjabiensis (Giraffidae, Ruminantia, Artiodactyla) from the Lehri Outcrops, Jhelum, Northern Pakistan. Pakistan Journal of Science 62, 120–123.
Khan MA, Ghaffar A, Ali A, Farooq U, Bhatti ZH, Akhtar M. 2005. Report on Mammalian Fossils of Chinji Formation, Dhulian, Pakistan. American Journal of Applied Sciences 2(12), 1619–1628.
Khan MA, Khan AM, Iqbal M. Akhtar M. 2009. Mammalian remains in the Chinji type locality of the Chinji Formation: a new collection, Journal of Animal and Plant Science 19(4), 224–229.
Lydekker R. 1876. Molar teeth and other remains of Mammalia from the India Terrtiaries. Pal. Indica 10(2), 12–87.
Matthew WD. 1929. Critical observations upon Siwalik mammals (exclusive of Proboscidea). Am. Mus. nat. Hist. Bull. 56, 437–560.
Meijaard E, Groves CP. 2004. A taxonomic revision of the Tragulus mouse deer (Artiodactyla). Zoological Journal of the Linnaean Society 140, 63– 102.
Meijaard E, Umilaela, Wijeyeratne G. 2010. Aquatic escape behaviour in mouse deer provides insight into tragulidevolution. Mammalian Biology 75, 471–473.
Milne−Edwards A. 1864. Recherches anatomiques, zoologiques et paléonto−logiques sur la famille des chevrotains. Annales de ScienceNaturelle Paris 5, 1– 167.
Morales J, Soria D, Sanchez IM, Quiralte V, Pickford M. 2003. Tragulidae from Arrisdrift, Basal Middle Miocene, Southern Namibia. Memoir Geological Survey Namibia 19, 359–370.
Nanda AC. 2008. Comments on the Pinjor Mammalian Fauna of the Siwalik Group in relation to the Post-Siwalik faunas of Peninsular India and Indo-Gangetic Plain. Quat. Int., 192, 6–13.
Pickford M, Senut B, Mourer-Chauvire C. 2004. Early Pliocene Tragulidae and peafowls in the Rift Valley, Kenya: Evidence for rainforest in East Africa. Comptes Rena’us Palevol 3, 179–189.
Pickford M, Senut B. 2000. Geology and Palaeobiology of the Namib Desert, southwestern Africa. Memoir of Geological Survey of Namibia 18, 1–155.
Pickford M. 2001. Africa’s smallest ruminant: a new tragulid from the Miocene of Kenya and the biostratigraphy of East African Tragulidae. Geobios 34(4), 437–447.
Pilgrim GE. 1937. Siwalik antelopes and oxen in the American Museum of Natural History. Bull. Amer. Mus. Nat. Hist. 72, 729–874.
Pilgrim GE. 1939. The fossil Bovidae of India. Pal. Indica., N .S. 26(1), 1–356.
Quiralte V, Sanchez IM, Morales J, Pickford M. 2008. Tragulidae (Artiodactyla, Ruminantia) from the Lower Miocene of the Sperrgebiet, Southern Namibia. Memoir of the Geological Survey of Namibia 20, 387–396.
Raymond LB, Kordos L, Rook L. 2004. Recent Advances on Multidisciplinary Research at Rudabánya, Late Miocene (MN9), Hungary: a compendium. Palaeontographia Italica, Pisa 89, 1–34.
Rossner GE. 2010. Systematics and palaeoecology of Ruminantia (Artiodactyla, Mammalia) from the Miocene of Sandelzhausen (southern Germany, Northern Alpine Foreland Basin). Paläontologische Zeitschrift 84, 123–162.
Samiullah K. 2011. Taxonomic studies of Fossil even and odd-toed Mammals from Miocene rocks of the Dhok Bun Ameer Khatoon, District Chakwal, Punjab, Pakistan. Ph.D. thesis, university of the Punjab, Lahore, Pakistan.
Sanchez IM, Quiralte V, Morales J, Pickford M. 2010. A new genus of tragulid ruminant from the early Miocene of Kenya. Acta Palaeontologica Polonica 55, 177-187. http://dx.doi.org/10.4202/app.2009.0087
Ullah K, Arif M, Shah MT. 2006. Petrography of Sandstones from the Kamlial and Chinji Formations, Southwestern Kohat Plateau, NW Pakistan. Impl. for Sour. Lithol. & Paleoecology 39, 1–13.
Vislobokova IA. 1997. Eocene–Early Miocene Ruminants in Asia, Mém. Trav. E.P.H.E., Inst. Montpellier 21, 215–223.
Vislobokova IA. 2001. Evolution and Classification of Tragulina (Ruminantia, Artiodactyla). Pal. J. Suppl. 35(2), 69–145.
Webb SD, Taylor BE. 1980. The phylogeny of hornless ruminants and a scription of the cranium of Archaomeryx. Bulletin of the American Museum of Natural History 167, 117–158.
Welcomme JL, Benammi M, Crochet JY, Marivaux L, Métais G, Antoine PO, Baloch I. 2001. Himalayan Forelands: palaeontological evidence for Oligocene detrital deposits in the Bugti Hills (Balochistan, Pakistan), Geol. Mag 138(4), 397–405.
West RM. 1980. A minute new species of Dorcatherium (Tragulidae, Mammalia) from the Chinji Formation near Daud Khel, Mianwali District, Pakistan. Contribution in Biology and Geology 33, 1–6.
Whitworth T. 1958. Miocene ruminants of East Africa. Fossil Mammals of Africa. Bulletin British Museum Natural History 15, 1–50.
Khizar Samiullah, Farhat Jabeen, Riffat Yasin, Sajid Yaqub, Fatima Jalal, Shehzad Ahmad, Bilal Rasool, Khurram Feroz, Muhammad Akhtar (2015), Dorcatherium from the dhok bun ameer khatoon, chinji formation, lower siwalik hills of Pakistan; JBES, V6, N3, March, P408-422
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